Despite the name, which suggests a version relatively light in feature changes over the previous version (like the transition from Leopard to Snow Leopard), Mountain Lion is intended to be a major new feature release that continues the work of bringing iOS features to the Mac: many of its major features are iOS transplants, including the Notification Center (which will bring unified notifications to OS X, replacing third-party apps like Growl), Game Center, iMessage support (in the form of an app called Messages, which replaces iChat - there's a free beta available for Lion users now), AirPlay Mirroring, a Notes app, Reminders, Twitter integration, tighter iCloud integration, and others. Frankly, this list of iOS imports actually seems to make more sense for the Mac as a platform than did some of the features (like Launchpad) that were brought over in Lion.
Mountain Lion will also include some new features all its own: Gatekeeper, which is aimed straight at system administrators, will allow admins to lock down the type of apps allowed to run on Macs. You can choose to allow apps only from the Mac App Store, apps from the Mac App store as well as those from developers you approve, or apps from anywhere (which is the default behavior in OS X currently). This can be seen as another step toward disallowing non-Mac App Store programs from running in OS X, but taken at face value it appears to be a solid compromise between the security of iOS-like behavior and the flexibility to install code from anywhere that users have always been accustomed to in OS X.
We don't have any information about system requirements yet, so we don't know whether Mountain Lion will run on any Lion-compatible Mac (which seems technically possible) or whether it will drop support for some older machines (which has historically happened with new OS X releases - see this page of our Lion review for in-depth information on what got dropped from the support list and why). The Apple developer site is currently down, but as soon as it comes back up those with developer accounts should be able to download and play with the next version of OS X. We'll continue to cover the new OS as details are made public.
Update: As we suggested might happen in our Lion review, Mountain Lion's developer preview appears to do away with support for any Mac that cannot boot into OS X's 64-bit kernel. I'll link you to that page of our Lion review again if you'd like deep technical information about what that means, but the short version is that a wide range of Apple's products from 2007 and 2008 are being dropped regardless of whether they include a Core 2 Duo processor. The list of supported Macs includes:
• iMac (mid 2007 or later)
• MacBook (13-inch Aluminum, 2008), (13-inch, Early 2009 or later)
• MacBook Pro (13-inch, Mid-2009 or later), (15-inch, 2.4/2.2 GHz), (17-inch, Late 2007 or later)
• MacBook Air (Late 2008 or later)
• Mac Mini (Early 2009 or later)
• Mac Pro (Early 2008 or later)
• Xserve (Early 2009)
The cutoff happens in different places for different products, but here are some rules of thumb: if your Mac uses the ATI Radeon X1600 graphics chip or the Intel GMA 950/X3100 integrated graphics chips, you're out of luck. If you've got a white iMac or one of the very first Mac Pros, you're out of luck. There are a few easy ways to check whether your Mac can run the 64-bit kernel, and Apple outlines all of them in this support document.
It should be noted that this information comes from the developer preview's release notes and may not be indicative of the final support list, but Lion's dropping of Core Duo Macs (and Snow Leopard's dropping of PPC Macs) were known quantities pretty early in the development of those operating systems - support for these older Macs may be added before the final release, but history suggests otherwise.
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